• Brown bears possess anal sacs and secretions may code for sex 

      Rosell, Frank; Jojola, Susan M.; Ingdal, Kristian; Lassen, Bård Andreas; Swenson, Jon; Arnemo, Jon Martin; Zedrosser, Andreas (Journal article; Peer reviewed, 2010-09-14)
      Olfactory communication occurs in carnivores and many scent-mark with anal gland secretions (AGS), which contain a variety of information including sex-related cues. Currently, there is disagreement about whether bear ...